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1、孟德爾遺傳定律本章重點(diǎn)1.孟德爾分離規(guī)律、驗(yàn)證、應(yīng)用;2.二對(duì)相對(duì)性狀的遺傳;3.多對(duì)相對(duì)性狀的遺傳;4.孟德爾自由組合規(guī)律、驗(yàn)證、應(yīng)用;5.遺傳學(xué)數(shù)據(jù)的統(tǒng)計(jì)處理第一節(jié)分離規(guī)律人類很早就從整體上認(rèn)識(shí)了遺傳現(xiàn)象:親子性狀相似?直觀上認(rèn)為子代所表現(xiàn)的性狀是父、母本性狀的混合遺傳,以后世代不再分離。Mendel’sExperimentsGregorJohannMendel(1822-84)Mendel,bornin1822,grewupinaruralareaofwhatisnowapartofCzechoslo
2、vakia.Becausehisfamilywaspoor,heenteredamonasteryinordertocontinuehiseducation.Mendelhadalifelonginterestinbiology,particularlyanimalbreeding,butbecausehissuperiorsdidnotthinkitappropriateforaclerictobreedanimals,heconcentratedinsteadonraisinggardenpeasand
3、otherplantsinthemonasterygarden.Mendel’sExperimentsTheresultsofhisworkwerepublishedinGermaninan1866paperentitled“experimentsinPlantHybridization.”Thepaperwasdistributedfairlywidelythroughlibraries,butMendel’scontemporariesdidnotunderstandhisfindings,probab
4、lypartlybecauseofthemathematicalexplanationsasteryin1868andpublishednothingfurtheronhereditybytherestofthescientificcommunityuntil1900,whenthreebotanists,CarlCorrensinGermany,HugodeVriesintheNetherlands,andErichvonTschermakinAustria,rediscoveredhisworkafte
5、reachhadapparentlyindependentlyreachedsimilarconclusions.孟德爾遺傳分析的方法嚴(yán)格選材從豆科植物中選擇了自花授粉且是閉花授粉的豌豆作為雜交試驗(yàn)的材料。精心設(shè)計(jì)采取單因子分析法。定量分析法首創(chuàng)了測(cè)交方法孟德爾試驗(yàn)的特點(diǎn):(1).遺傳純合:以嚴(yán)格自花授粉植物豌豆為材料;(2).穩(wěn)定性狀:選擇簡(jiǎn)單而區(qū)分明顯的7對(duì)性狀進(jìn)行雜交試驗(yàn)研究;(3).相對(duì)性狀:采用各對(duì)性狀上相對(duì)不同的品種為親本;(4).雜交:進(jìn)行系統(tǒng)的遺傳雜交試驗(yàn);(5).統(tǒng)計(jì)分析:系統(tǒng)記載各世
6、代中不同性狀個(gè)體數(shù),應(yīng)用統(tǒng)計(jì)方法處理數(shù)據(jù),獲得各種結(jié)果,否定了長(zhǎng)期流行的混合遺傳觀念。WhyMendelchosethepeaplant?Thegardenpeawaseasytocultivateandhadrelativelyshortlifecycle.Theplanthaddiscontinuouscharacteristicssuchasflowercolorandpeatexture.Inpartbecauseofitsanatomy,pollinationoftheplantcouldbeco
7、ntrolledeasily.Foreignpollencouldbekeptout,andcross-fertilizationcouldbeaccomplishedartificially.TheanatomyandlifecycleofthepeaplantArtificialcrossofawhite-flowered(maleparent)withaviolet-flowered(femaleparent)peaplant孟德爾認(rèn)為父母本性狀遺傳不是混合,而是相對(duì)獨(dú)立地傳給后代,后代還會(huì)分離出父
8、母本性狀。 提出:①.分離規(guī)律;②.獨(dú)立分配規(guī)律。1.1.1Concernedconcept性狀(trait):生物體所表現(xiàn)的形態(tài)特征和生理特性,并能從親代遺傳給子代?!? (1)單位性狀(unittrait):個(gè)體表現(xiàn)的性狀總體區(qū)分為各個(gè)單位之后的性狀。例如:豌豆的花色、種子形狀、株高、子葉顏色、豆莢形狀及豆莢顏色(未成熟)。 (2)相對(duì)性狀(contrastingtrait):指同一單位性狀的相對(duì)差